I won't proclaim to be a biologist, because I most definitely am not, but from my own nerding out, recently watching a bunch of mycology documentaries, it's not so much that the root systems themselves are grafted or physically connected, but that an entire ecosystem of fungal roots and what-not share nutrients from root system to root system in a rather impressive display of symbiosis.
My expertise is fire life safety, so YMMV in the above, but it certainly made sense to me (and was pretty cool to learn, to boot)